Education

• Ph.D. Evolutionary Biology. Committee on Evolutionary Biology. The University of Chicago. (2021)
• M.Sc. Biology. University of Puerto Rico, Río Piedras Campus (2011)
• B.A. Environmental Sciences. University of Puerto Rico, Río Piedras Campus (2002)

Other training

• February 17 to 21, 2014. International Lichen Field Symposium, Mata de Plátano Field Station, Puerto Rico. (Organizer and Participant)
• May 21 - June 1, 2012. Tropical Lichens and Forest Health: A Multidisciplinary Approach. Organization for Tropical Studies (OTS), Costa Rica.
• October 3-7, 2011. Taller de Liquenología Tropical: Los Micro- y Macroliquenes de Puerto Rico. Universidad de Puerto Rico, Recinto de Rio Piedras. (Organizer and Participant)
• March, 2009: 18th Annual Tuckerman Workshop, Florida, USA. (http://www.abls.org/) (Participant)
• September, 2005: ‘Liking Lichens.’ Exploring Lichen Ecology and the Environment: An Educator Enhancement Initiative to Train Teachers and Build a State-wide Bio-Monitoring Project. Augusta, Georgia. (http://www.uga.edu/crms/lichens.htm) (Co-Instructor and Participant)
• October, 2004: International Workshop of Tropical Lichenology (Ticolichen: The Costa Rican Lichen Biodiversity Inventory: http://www.fieldmuseum.org/research_collections/botany/botany_sites/ticolichen/index.html) (Participant)

Organizations

• Society for the Study of Evolution
• Ecological Society of America (ESA)
• American Bryological and Lichenological Society (ABLS)
• International Association of Lichenology (IAL)
• Mycological Society of America (MSA)

Growth forms of different lichens from Puerto Rico. (left) Crustose – Thalloloma rubromarginatum from El Yunque National Forest (EYNF); (center) Foliose – Sticta tainorum from Bosque Estatal de Toro Negro, central Puerto Rico; (right) Fruticose – Cladonia robusta from Reserva Natural Laguna Tortuguero, northern Puerto Rico.